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ENCYCLOPEDIA OF EARTH AND ATMOSPHERIC SCIENCES ENCYCLOPEDIA OF MATHEMATICAL SCIENCES ENCYCLOPEDIA OF BIOLOGICAL,PHYSIOLOGICAL AND HEALTH SCIENCES ENCYCLOPEDIA OF SOCIAL SCIENCES AND HUMANITIES ENCYCLOPEDIA OF PHYSICAL SCIENCES,ENGINEERING AND TECHNOLOGY RESOURCES ENCYCLOPEDIA OF CHEMICAL SCIENCES,ENGINEERING AND TECHNOLOGY RESOURCES ENCYCLOPEDIA OF WATER SCIENCES,ENGINEERING AND TECHNOLOGY RESOURCES ENCYCLOPEDIA OF ENERGY SCIENCES,ENGINEERING AND TECHNOLOGY RESOURCES ENCYCLOPEDIA OF ENVIRONMENTAL AND ECOLOGICAL SCIENCES,ENGINEERING AND TECHNOLOGY RESOURCES ENCYCLOPEDIA OF FOOD AND AGRICULTURAL SCIENCES,ENGINEERING AND TECHNOLOGY RESOURCES ENCYCLOPEDIA OF HUMAN RESOURCES POLICY AND MANAGEMENT ENCYCLOPEDIA OF NATURAL RESOURCES POLICY AND MANAGEMENT ENCYCLOPEDIA OF DEVELOPMENT AND ECONOMIC RESOURCES ENCYCLOPEDIA OF INSTITUTIONAL AND INFRASTRUCTURAL RESOURCES ENCYCLOPEDIA OF TECHNOLOGY,INFORMATION, AND SYSTEMS MANAGEMENT RESOURCES ENCYCLOPEDIA OF REGIONAL SUSTAINABLE DEVELOPMENT REVIEWS

The above simplified figure illuminates the essential interconnectedness of the sixteen component encyclopedias of EOLSS.

 In the real world, the various knowledge domains do not exist in isolation from each other. They form an integrated whole, with links in all directions. It is well known that all forms of human knowledge are inter-connected and inter-related. EOLSS mimics this complexity, the automatic inter-connectedness of the various subject categories facilitating navigation through the vast landscape of EOLSS knowledge. This provides the user with an effective and efficient tool to search, navigate and browse through each of the component encyclopedias, through any combination of the sixteen, or through the whole of EOLSS.

ENCYCLOPEDIA OF BIOLOGICAL, PHYSIOLOGICAL AND HEALTH SCIENCES

CONTENT OUTLINE (partial listing)

 

BIOLOGICAL SCIENCE FOUNDATIONS

    The Beginning of Modern Biological Science

        What Is Biological Life?

        Microscopic Life

        The Origin of Life

        Evolution and the Origin of Species

    Genetics and Evolution

        Genetics in the Modern World

        The Mechanisms Involved in Evolution

        Humanity, the Environment, and Evolution

        Artificial Evolution

    The Molecular Basis of Life

        The Cell

        The Gene

        The Biochemical Processes of Life

        Genetic Manipulation

    The Molecular Tree of Life

        Morphological Methods of Classifying Species

        Biochemical Methods of Classifying Species

        Gene-Based Classification

        Life On Earth Over the Last 4 Billion Years

    The Limits of Life on Earth

        Water as a Limiting Factor

        Ionic Concentration as a Limiting Factor

        Temperature as a Limiting Factor

        Acidity/Alkalinity as a Limiting Factor

        Gaseous Atmosphere as a Limiting Factor

        Energy as a Limiting Factor

        The Edges of the Envelope

    Life Elsewhere in the Universe

        Is Carbon-Based Life Alone?

        Carbon-Based Life on the Planets

        Carbon-Based Life in Other Star Systems

        Panspermia

 

ORGANIC CHEMICALS INVOLVED IN LIFE PROCESSES

    Proteins

        Amino Acids

        Polypeptide chains

        Primary structure

        Secondary structure

        Tertiary structure

        Quaternary structure

        Enzymes

    Carbohydrates

        Monosaccarides

            Stereo-isomerization

            Cyclic Structures

        Disaccharides

    Polysaccharides

    Lipids

        Fatty acids

        Fatty acids in humans

        Triacyglycerols

        Phospholipids

        Steroids

    Nucleic Acids

        Nucleotides

        The nucleic acids

            DNA

            DNA can have different structural features

            RNA

 

CARBON FIXATION

Carbon fixation in higher plants

The Light Dependent Reactions of Plant Photosynthesis

The Light Independent Reactions of Photosynthesis: Synthesis of Carbohydrates

C3, C4 and CAM metabolism

Algae

Bacteria

Purple Bacteria

Green Sulfur Bacteria

Green Gliding Bacteria

Heliobacteria

Global photosynthesis and the atmosphere

 

EUKARYOTE CELL BIOLOGY

    Origin of Eukaryotes

    Cellular differentiation in multicellular organisms

        Plants

        Animals

    Eukaryotic cell structure

    Organization of eukaryotic cells

        Plasma membrane

        Extracellular matrices

        Proteins synthesis and transport

        Cytoskeleton and movement

        Nucleus

            Genomes

            Gene expression

            Maintaining the genome

        Organelles

    The cell cycle

        Mitosis

        Meiosis

    Regulation of cell growth

        Signal transduction

        Programmed cell death

        Cancer

    Experimental models

        Yeast

        Arabidopsis

        Drosophila

        The mouse

        Cell culture

        Separation of cellular contents

        Tracing biochemical pathways

    Future Investigations

 

CELL THEORY, PROPERTIES OF CELLS AND THEIR DIVERSITY

    The Composition of Life

        Origin of organic molecules

        RNA and the origin of life

    Cell as the unit of life

        Cell theory

        The theory of organisms

        Cell size

    The diversity of life

        Origin of the first cell

        The endosymbiont hypothesis

    Cellular diversity

        Plant tissue diversity

            Dermal tissue

            Vascular tissue

            Ground tissue

        Animal tissue diversity

            Epithelial tissue

            Muscle tissue

                Connective tissue

                Nervous tissue

                Blood

                Sensory cells

                Germ cells

    Tissue maintenance and renewal

        Tissue renewal rates

        Stem cells

    Discussion

 

CELL MORPHOLOGY AND ORGANIZATION

    The cytoplasm

    Membranes

        Structure

        The cell cortex

        Transmembrane proteins

        Endoplasmic reticulum

        Golgi apparatus

        Lysosomes

    Extracellular matrices

        Glycocalyx

        Cell walls

    Cytoskeleton

        Microfilaments or Actin filaments

        Intermediate filaments

        Microtubules

    Cilia and Flagella

    Connections between cells

         Plasmodesmata

         Desmosomes

         Tight junctions

         Gap junctions

    Vacuoles

        Central vacuoles

        Contractile vacuoles

    Cell organization

    Membrane transport systems

        Passive transport

        Active transport

        Endocytosis

    Protein synthesis

        Initiation

        Elongation

        Termination

    Protein targeting

        Proteins made in the cytoplasm

        Chaperones

        Proteins made on the rough endoplasmic reticulum

        The Golgi apparatus

        Vesicles

    Protein breakdown

        Proteosomes

        Lysosomes

    Cell movement

        Microfilaments and cell movement

        Myosin

        Microtubules

        Cilia and flagella movement

 

CELL NUCLEUS AND CHROMATIN STRUCTURE

    The nucleus

        The nuclear membrane

        The nuclear pore

            Structure of the nuclear pore

            Function of the nuclear pore

        The lamina

    The genome

    Chromosomes

        The nucleosome

            Histones

            DNA condensation

        Functional domains

            Centromere

            Telomere

    DNA replication

        Origin of replication

        DNA polymerases

        Nucleosome replication

    Nucleolus

        rRNA genes and expression

        Ribosome assembly and export

    Transcription

        RNA polymerases

        Eukaryote promotors

        Initiation of transcription

        RNA elongation

        Termination of transcription

        Regulation of transcription

        Chromatin structure and transcription

    mRNA processing and turnover

        Capping

        Polyadenylation

        Splicing

        RNA editing

    tRNA processing and turnover

 

ORGANELLES AND OTHER STRUCTURES IN CELL BIOLOGY

The distribution and function of the mitochondrion

The structure of the mitochondrion

The mitochondrial genome

The mitochondrial genetic code and translation mechanism

Maternal inheritance of the mitochondrion

Distribution and structure of the chloroplast

The chloroplast genome

The chloroplast genetic code

Maternal inheritance of the chloroplast

An endosymbiotic origin for the mitochondrion and chloroplast

Evolution of the mitochondrion

Evolution of the Chloroplast

Other possibly endosymbiotic organelles

 

MITOSIS, CYTOKINESIS, MEIOSIS AND APOPTOSIS

    The eukaryote cell cycle

        Phases

    Mitosis

        Prophase

        Metaphase

        Anaphase

        Telophase

        Cytokinesis

    Meiosis

        Stages of meiosis

    Fertilization and development

    Regulators of Cell cycle

        Checkpoints

            G1/S checkpoint

            G2/M checkpoint

        Mitosis checkpoint

        Maturation promoting factor

        Cyclin dependent protein kinases

            Diversity and action

            Regulation

            Cyclin regulation of mitosis

        Growth factors

        Inhibitors of cell cycle progression

    Programmed cell death

        Triggers of apoptosis

        Pathways leading to apoptosis

 

CELL GROWTH REGULATION, TRANSFORMATION AND METASTASES

    Signal molecules

        Transmembrane signals for large signal molecules

        Small signal molecules

    Switches of intracellular molecules

        Regulation of signal molecules

    Cell surface receptors

         Ion-channel-linked receptors

        G-protein-linked receptors

            General structure and function of G-protein-receptors.

            G-proteins and ion-channels

            G-proteins and enzymes.

        Enzyme-linked receptors

            Protein tyrosine kinases

            Signaling domains SH2 and SH3.

            Ras/MAPK pathway

        The JAK/STAT pathway

    Cancer

        Ras proto-oncogenesis

        Signal transduction

        Kinases

        Insertions, deletions and translocations

        Viral oncogenesis

            DNA viruses

            RNA viruses

        Tumor suppressor genes

 

MICROBIOLOGY

Taxonomy

Morphology

Eucaryote Microorganisms - Protozoa

Eucaryote Microorganisms - Algae

Eucaryotic Microorganisms - Fungi

Procaryotic Microorganisms – Bacteria and Archaea

Viruses

Chemical Taxonomy

Molecular Taxonomy

Protein based taxonomy

DNA based taxonomy

Ribosomal DNA based taxonomy

Limitation of single gene based taxonomy

Genome based phylogeny and taxonomy

Genome sequencing

Genomics and phylogeny

The future for genome phylogenies

Horizontal genetic exchange

Transformation

Transduction

Conjugation

Procaryote and Eucaryote Microbial Cell Structure

Procaryote Cell Structure

Bacterial Spores and Cysts

The subcellular structure of eucaryotic Microorganisms

Morphology of Fungi

Morphology of Protists

Cultivation of Microorganisms

Control of Microorganisms

Major Groups of Procaryotes

Major Groups of Eucaryotic Microorganisms

Viruses

Pathogenesis and Microorganisms

Antibiotics and Microorganisms

Microbial Biotechnology

 

PROKARYOTIC CELL STRUCTURE AND FUNCTION

    Nucleotide

    Cytoplasmic Matrix

        Ribosomes

        Plasmids, episomes and bacteriophage

        Inclusion bodies and storage granules

        Endospores

        Intracytoplasmic membranes

    The Cell Envelope

        Cytoplasmic Membrane

        The Periplasmic Space

        The Cell Wall

        The Outer Membrane

    Components Exterior to the Cell Wall

        EPS

        Flagella

        Pili

        Fimbriae

    Differentiation and multicellularity

        Differentiation in Actinomycetes

        Multicellularity and differentiation in cyanobacteria

        Differentiation in Myxobacteria

 

PROKARYOTIC DIVERSITY

    The Archae, Cyanobacteria, Green phototrophs, and Deeply Branching Genera

        Thermoprotei, Sulfolobi and Barophiles

        The Methanogens

        The Halobacteria

        The Thermoplasms

        The Thermococci

        Aquifex and relatives

        Thermotogas and Geotogas

        The Deinococci

        Thermi

        Chrysiogenes

        The Chlorflexi and Herpetosiphons

        Thermomicrobia

        Prochloron and the Cyanobacteria

        Chlorobia

    Proteobacteria

        The a -Proteobacteria

        The b Proteobacteria

        The c Proteobacteria

        The d Proteobacteria

        The e Proteobacteria

    The low G+C gram-positives

    The high G+C gram-positives

    The Planctomycetes, Spirochetes, Fibrobacter, Bacteroides and Fusobacteria

        The Planctomycetes

        The Spirochetes

        The Fibrobacters

        The Bacteroides

        The Flavobacteria

        The Sphingobacteria, Flexibacteria and Cytophaga

        The Fusobacteria

 

PROKARYOTE GENETICS

    Mechanism of DNA Mutation, Transfer and Recombination in Bacteria

    Mutation

        Types of mutations

        DNA damage

        DNA Repair

    Transformation

        Gram -ve Bacteria

        Low G+C% Gram +ve Bacteria

        High G+C% Gram +ve Bacteria

        Archaea

    Transduction

        Generalized Transduction

        Specialized Transduction

        Importance of Generalized and Specialized Transduction.

    Conjugation

        Plasmids

            Plasmids in Gram -ve Bacteria

            Plasmids in Low G+C% Gram +ve Bacteria

            Linear Plasmids

            Plasmids in the Archaea

        Conjugation

            Gram -ve Bacteria

            Low G+C% Gram +ve Bacteria

            Linear Chromosomes

    Recombination in viruses

    Prokaryote Genetics and Evolution

        Multiple Drug Resistant Plasmids

        Transposition

        Gene Scavenging

        Bacterial Species, do they really exist?

 

PROKARYOTIC GROWTH, NUTRITION AND PHYSIOLOGY

    Bacterial Cell Growth and Division

    The bacterial cell cycle and its regulation

    Bacterial Population Growth

    Bacterial Nutrition

        Carbon sources and assimilation

        Nitrogen sources and assimilation

        Phosphate sources and assimilation  

        Sulfur sources and assimilation

        Nutrient reserves

    Energy generation

        Aerobic respiration

        Anaerobic respiration

        Fermentation

        Photosynthesis

    Bacterial Nutrient Stress Responses

 

BIOLOGICAL SCIENCE FUNDAMENTALS (SYSTEMATICS)

    Life on Earth

    The Geological Scenario and the Major Evolutionary Transitions

        Some Fundamentals of Geology

        Geological Changes, Evolutionary Transitions, and Extinctions

            Major Evolutionary Transitions

    The Cell

        The Procaryotic Cell

        Autotrophs and Heterotrophs

        The Eucaryotic Cell

        The Emergence of Eucaryotes and the Kingdoms of Life

    Routes to Multicellularity

        Aggregative

        Plasmodial

        Clonal

        Soma and Germ Plasm

    Growth and Development

    Life-Cycles

        Direct and Indirect Development

        The Life-Cycles of Cnidarians, Rotifers, and Aphids

        Plant Life-Cycles

    Individual, Colony, Society

    Populations, Species, and Communities

    The Continuity of Life

        Descent with Modification

        Mutations

        Gene Assortment: Inheriting Variation

        Selection, Random Change

    Adaptation

    Life Forms

        Living In the Sea

        Nutrition

        From the Sea to the Internal Waters

        Water to Land

    Biodiversity

    The Science of Taxonomy

 

HISTORY AND SCOPE OF BIOLOGICAL SCIENCES

    Ancient and Medieval Times up to the 16th Century

    Post-Renaissance Developments

    Paleontology and Evolution

    Morphology and Physiology

    Genetics

    Behavior

    Ecology and Applied Ecology

 

HISTORY OF BIOLOGY

    Antiquity

    Medieval and Renaissance times

    The development of morphology

    Palaeontology

    Taxonomy and Evolution

    Histology, Reproduction and embryology

    Physiology

    Genetics

    Ecology and Ethology

    Pathology

 

CHARACTERISTICS OF LIVING BEINGS

    Former Conceptions of Life

        The Birth of Biology

        Metabolism, 'Irritability' and Cellular Organisation

        The Becoming of Individuals and Species  

        Inadequacy of Mechanistic Interpretations and the Revival of Vitalism

        The Gestation of the Concept of Biological Information

        Information Flux is an Endowment of Every Living Being

        First Conclusions on Living Beings

    Current Conceptions About Living Beings

        The Rise of Genetics

        The Mighty Advance of Dynamic Biochemistry

        The Sources of Information Flux and the Self-Control in Living Beings

        Limits and Peculiarities of the Sensory Flux

        Self Regulation of Genetic Information

        Information and Needs

        Death and Life

        Ontogeny and Morphogenesis

    Evolution

        Is it Conceivable that a Living Being May Not Evolve?

       The Darwinian Assumptions

       The Subjects of Darwin's Evolution

    Summary of the Properties of Living Beings

 

LEVELS OF BIOTIC ORGANISATION

    The development of organisation at individual level

    The development of organisation: embryology and cycles

    The palaeontological account

    The intertaxa organisation and evolution

    Symbioses and parasitism

 

POPULATIONS, SPECIES AND COMMUNITIES

    Populations and Species

       The Species Concept

       The Population Concept

           Population Size

           Constant Reproduction of Populations

           Variation in Population Size

    Species Communities

        General Aspects

        Composition of Communities

        Island Biogeography

        Biological Invasions

        Interaction

            Interspecific Competition

            Mutualism

            Host-parasite Relations

            Predator-prey Relations

 

PHILOSOPHY OF BIOLOGICAL SCIENCES

Basic issues in the philosophy of biology: Is biology an experimental or an historical discipline?

    Methodological debates

    Realism versus nominalism

    The population concept: problems and inferences

    Bioethics: some sample problems

 

THE ORIGIN AND EVOLUTION OF EARLY LIFE

    The need for organic molecules

    Asymmetry in organic molecules

    The need for suitable energy

    Rise and fall of organic chemistry of galaxies

    Organic chemistry on orbiting bodies

    The need for liquid water

    Evolution of hydrospheres

    Self-organization and self-duplication

    Encapsulation and translation

    The first ecosystems

    Probability and stability of a biosphere

 

THE BUILDING BLOCKS OF PRIMITIVE LIFE: ORIGIN AND FORMATION

    The role of water

    Possible environments for the production of prebiotic organic molecules

        Production of CHONS in the atmosphere

        Submarine hydrothermal systems

        Delivery of extraterrestrial CHONS

    Availability of the primitive building blocks

    Production of homochiral building blocks

 

FROM THE BUILDING BLOCKS TO LIFE

    A Primitive Cellular Life

        Prebiotic RNA

            The Montmorillonite Clay-catalyzed Synthesis of RNA

            Metal Ion Catalysis of RNA Synthesis

            Template-directed RNA Synthesis

        Prebiotic Polypeptides

            Polypeptide Formation on Clay Minerals

            Peptide Elongation on Minerals

            Polypeptide Formation via N-Carboxyanhydrides

            Proteinoids

            Polypeptide Formation Using Simulated Hydrothermal Conditions

            Thermal Conversion of Amino Acid Amides to Polypeptides

            Prebiotic Polypeptide Structure and Stability

            Prebiotic Polypeptide Catalysis

    Primitive Life Based on RNA: The RNA World

        RNA Analogs and Surrogates

            Nucleotides with a Pyrophosphate Backbone

            Pyranosyl-RNA, a Hexose Nucleic Acid, and Threose-RNA, a Tetrose Nucleic Acid

            Peptide Nucleic Acid

    Autocatalysis Preceding RNA

 

THE EARLIEST ANAEROBIC LIFE AND EARLY AEROBIC LIFE

    The first cells

    From anaerobic heterotrophy to autotrophy

    Ancient oxygen-producing photosynthesis

    The first aerobic microorganisms

    Origin of organelles of bacterial cells

    Origin of the nucleus

    Origin and evolution of the mitochondrion

    Origin and evolution of plastids

    Origin of the cilium

    Rise of highly differentiated organisms

 

EVOLUTION

    Major Transitions in Evolution

        Natural Selection and Levels of Organization

    Different Approaches to the Study of Evolution

    Microevolution

        The Causes of Evolution

        Natural Selection

    Adaptive Evolution

        Adaptation

        Adaptations and Adaptive Responses

        Adaptation and Exaptation

        Coevolution

    Neutral Evolution

    Species and Speciation

    Macroevolution

    Phylogeny

    Evolution, Complexity, and the Information Content of Living Beings

 

HISTORY OF EVOLUTIONARY THEORY

    Scientific Historiography and the Analysis of the Theoretical Concepts

    The Definition of "Species"

    The Problem of Classification

    Linnaeus, the "System" and the "Methods"

    Buffon and the nuance of Nature

    The Historical Dimension and the Re-orientation of Natural History

    Lamarck and Transformism

    Paleontology and Geology

 

THE DARWINIAN VIEW OF LIFE

    Evolutionism and Darwinism

    "What are the Laws of Life?"

    The Structure of a New Theoretical Picture

    The Origin of Species and its Critics

    Toward a Natural History of Mind

 

EVOLUTION AND THE SPECIES CONCEPT

    Historical Aspects

        Pre-Darwin

        Darwin's View

        Post-Darwin (Modern Synthesis)

    Current Paradigms and Challenges

        Philosophical Perspectives: The Individual Species Concept

        Current Paradigm: The Biological Species Concept

            Isolating Barriers

            Limitations of the Biological Species Concept

                Non-universality

                Inapplicability to Multi-dimensional Situations

                Incomplete Speciation

        Challenges, Extensions and Alternatives to the Biological Species Concept

            Evolutionary Species Concepts

            Phenetic Species Concept

            Ecological Species Concept

            Recognition Species Concept

            Cohesion Species Concept

            Phylogenetic Species Concepts

    A Case Study

    The Future: Towards a Holistic Approach?

        Which Criteria?

        Monism versus Pluralism

 

SPECIATION AND INTRA-SPECIFIC TAXA

Time frame for speciation

Ecological considerations

Genetic models of speciation

Speciation by natural selection

Wright’s shifting balance theory

Founder-effect models

How many "speciation genes"?

Genetic properties of species

Modes of speciation

Allopatric speciation

Intra-specific taxa

Clinal variation

Vicariance model

Peripatric model

Reinforcement

Parapatric speciation

Stasipatric speciation: the role of chromosomes

Chromosome rearrangements and meiotic segregation

Post-zygotic isolation: two case studies

Sympatric speciation

Speciation in plants

Speciation by polyploidy

Speciation by hybridization

 

NATURAL SELECTION AND THE EFFECTS OF ECOLOGICAL INTERACTION ON POPULATIONS

    Natural Selection

        The Different Aspects of Natural Selection

            Stabilising Selection

            Directional Selection

            Disruptive Selection

            Frequency-dependent Selection

            Density-dependent Selection

            Sexual Selection

            Kin Selection

            Indirect Selection

    Non-selective Aspects of Population Biology

        Genetic Drift

            Founder Effect

            Bottleneck Effect

            Gene Flow

    Deterministic and Stochastic Concerns in Population Biology

        Interactions between Genotypes and Environment

        Metapopulations

            Local Extinction

            Recolonisation

 

CELL AND TISSUE STRUCTURE IN ANIMALS AND PLANTS

    Cell and tissue development

        Development of Animal Cells and Tissues

        Developmental Processes in Higher Plants

            Tissue Origins in Higher Flowering Plants

            Cell Division, Growth and Differentiation

            Initials in Meristems

        The Size of Cells

        Methods in Histology and Cytology

    Cell Ultrastructure

        Cell Membrane

        Plant Cell Wall

            Cellulose and Other Cell Wall Substances

            Cell Wall Formation

            Middle Lamella, Primary Cell Wall and Secondary Cell Wall

            Communication between Cells

        Cytoplasm

        The Nucleus of Eukaryotic Cells

            Nuclear Membrane

            Genetic Material

        Other Structures in the Cytoplasm

            Mitochondria

            Plastids

            Endomembrane System (Lysosomes, Golgi Complex, ER, Vacuoles)

            Ribosomes

            Cytoskeleton (Microtubules, Actin Filaments)

            Projections from the Animal Cell Surface (Flagella, Cilia, Microvilli)

    Selected Examples of Differentiated Cell Types

        Pigment Cells

        Muscle Cells

        Plant Root Hairs

        Vessel Elements in Plants

    General Organisation of Tissues

        Tissue Systems in Plants

        Animal Extracellular Matrix (ECM)

    Some Examples of Specialised Tissue Types

        Plant Epidermis

        Xylem

        Animal Cartilage and Bone

        Epithelia

        Nervous Tissue

 

BIOLOGICAL HOMEOSTASIS

    Stabilization

    Homeostasis

        Logic and Structure of the Homeostat

        The Components of the Homeostat: the Measuring Unit

        The Components of the Homeostat: The Effector

        The Control of Controllers

        The Control of Stores

    Homeorrhesis, the Control of Behavior

        Trajectory Control

        Smooth Control

        Coordinate Control in Homeostasis and Homeorrhesis

    Control of Development and Reproduction

        Exogenous and Endogenous Clock and the Calendar

        Embryonic Regulation

        Control of the Flux of Genetic Information

 

PATTERNS AND RATES OF SPECIES EVOLUTION

    Biological and morphological species concepts

    Evidence from ecology and palaeobiology

        The cichlid fishes of the African rift valley

        Sticklebacks and speciation by natural selection

        Ice ages and species migrations

       The punctuated equilibrium revolution

        Stasis

        Speciation and ecophenotypic change: the rift valley snails

    Fossil evidence of speciation.

        Microfossils

        Shallow marine invertebrates

        Terrestrial vertebrates

    Species diversification through time

        Biodiversity today

        Biodiversity through time patterns of increase

        Biodiversity through time land and sea compared

        Explanations for patterns of diversification

    Evolutionary patterns and processes

 

CLASSIFICATION AND DIVERSITY OF LIFE FORMS

    Natural vs. artificial classifications

    Classification vs. system

    Nomenclature

        The international codes of nomenclature

        Nomenclatural stability: problems and solutions

        Ethnotaxonomy

    The diversity of Life

        Nature and origin of biological diversity

        Species known and estimated

        Big genera

    Taxonomy and nomenclature of domesticated animals and cultivated plants

    The tools of classification

 

HISTORICAL REVIEW OF SYSTEMATIC BIOLOGY AND NOMENCLATURE

    The Origins

    From Classical Antiquity to the Renaissance Encyclopaedias

    From the First Monographers to Linnaeus

    Concepts and Definitions: Species, Homology, Analogy

    The Impact of Evolutionary Theory

    The Last Few Decades

    Nomenclature

    Natural History Collections

 

METABOLIC DIVERSITY IN PROKARYOTES AND EUKARYOTES

    The Thermodynamic and Mechanistic Basis of Cellular Metabolism

        Dissimilatory Metabolism

            The Phototrophic Way of Life

            The Chemheterotrophic Way of Life

            The Chemoautotrophic Way of Life

        Assimilatory Metabolism

            Autotrophy

            Heterotrophy

    Metabolic Diversity Within the Three Domains of Life: Archaea, Bacteria and Eucarya

    Phototrophy The Use of Light as an Energy Source

        Oxygenic Photosynthesis of Plants and Cyanobacteria

        Anoxygenic Photosynthesis in Bacteria

        Retinal Pigment-Based Phototrophy

    Heterotrophy

        Aerobic Respiration

        Anaerobic Respiration

        Fermentation

    Chemoautotrophy

        Nitrification

        Oxidation of Reduced Sulfur Compounds

        Oxidation of Reduced Iron and Manganese

        Aerobic Oxidation of Hydrogen

        Anaerobic Oxidation of Hydrogen: Methanogens and Homoacetogens

    Metabolic Diversity and the Cycles of Carbon, Nitrogen, Sulfur and Other Elements

 

DIVERSITY OF FORM, FUNCTION AND ADAPTATION IN MICROORGANISMS

    Cell Size and Cell Shape in the Procaryotes

        Small is Powerful

        Appendages and Other Strategies to Increase Cell Surface

        The Largest Procaryotes

    The Outer Layers of the Procaryotic Cell

        The Cell Wall and the Cytoplasmic Membrane

        Fimbriae and Pili

        Exopolysaccharides

    Special Intracellular Structures

        Intracellular Membrane Structures

        Storage Materials

    Motility Organelles

        Flagella

        Magnetosomes

        Gas Vesicles

    Survival and Dispersal Forms

    Multicellular Differentiation and Special Life Cycles in Procaryotes

        Heterocysts and Other Specialized Cells of Cyanobacteria

        Bdellovibrio, the Intracellular Predator of Bacteria

        The Life Cycle of Caulobacter and Hyphomicrobium

        The Life Cycle of Myxobacteria

Diversity of Form, Function, and Adaptation The Lower Eucaryotes and the Procaryotes Compared

 

DIVERSITY OF FORM, FUNCTION, AND ADAPTATION IN FUNGI

    Diversity of Form

        Hyphae and Mycelium

        Fungal Cells

        Colonization of Substrates

    Reproduction

        Sexual Reproduction

        Asexual Reproduction

        Spore Dispersal

    Diversity of Function

        Nutrition

        Saprotrophic Fungi

        Parasitic Fungi

        Symbiotic Fungi

            Mycorrhizas

            Lichens

            Endophytes

    Adaptation

        Adaptation to Terrestrial Habitats

        Temperature

    Economic Relevance of Fungi

        Plant diseases

        Food Spoilage

        Mycotoxins

        Drugs

        Food Production

        Edible Mushrooms

 

EVOLUTIONARY AND MOLECULAR TAXONOMY

    Basic Concepts

        Metabolism, Proteins, and Nucleic Acids

        Nuclear and Organellar Genomes

        Inheritance of Genomes

        Mutation and Molecular Evolution

    Molecular Methods used in Systematics

        Allozymes

        DNA/DNA Hybridization

        Restriction Fragment Length Polymorphisms (RFLP)

        Restriction Site Mapping

        DNA Sequencing

    Phylogenetic Analysis

        Distance Methods

        Parsimony Methods

        Maximum Likelihood Methods

        Relative Merits of the Three Approaches

        Reliability of Inferred Trees

        Questions and Precautions

    Future Developments

 

SYSTEMATICS OF THE MICROBIAL KINGDOM(S) AND FUNGI

    The Emergence of the Microbial World

        Timetable of Events

        Universal Ancestor and Early Eucaryotes

    Microbial Systematics

        Higher Order Classification

        Different Approaches to the Study of Microbial Diversity

            The Study of Procaryotes

            The Study of Eucaryotes

    Microbial Species Concepts

        Eucaryotic Species

        Procaryotic Species

    Rules of Nomenclature

    The Microbial Tree of Life

        The Procaryotes

        The Eucaryotes

 

SYSTEMATICS OF ARCHAEA AND BACTERIA

    Systematics, Taxonomy and Nomenclature of Procaryotes A Few Definitions

    Procaryote Systematics - A Historical Overview

    The Formal Framework of Description and Nomenclature of Procaryotes

    Approaches to the Classification of Procaryotes

        Experimental Tools of Procaryote Taxonomy

        The Species Concept in the Procaryote World

        The Number of Described Procaryote Species

    Archaea and Bacteria, the two Domains of the Procaryotic World

        The Recognition of the Archaea as a Separate Domain

        The Differences between Archaea and Bacteria

        Systematics of Archaea and the Properties of the Main Groups of Archaea

        Systematics of Bacteria and the Properties of the Main Groups of Bacteria

    How Many Procaryote Species are there in Nature?

 

PROTOCTISTA

    Classification

        Archamoeba

        Neomonada 

        Rhizopoda

        Mycetozoa

        Foraminifera

        Heliozoa

        Radiozoa

        Percolozoa

        Euglenozoa

        Dinozoa 

        Metamonada

        Parabasala

        Apicomplexa

        Ciliophora

 

SYSTEMATICS OF FUNGI

    Kingdom Fungi

        Phylum Chytridiomycota

        Phylum Zygomycota

            Class Zygomycetes

            Class Trichomycetes

        Phylum Ascomycota

            Class Archiascomycetes

            Class Saccharomycetes

            Class Ascomycetes

                Group Plectomycetes

                Group Pyrenomycetes

                Group Discomycetes

                Group Loculoascomycetes

                Lichenized Ascomycetes

        Conidial or Mitosporic Fungi

        Phylum Basidiomycota

            Class Urediniomycetes

            Class Ustilaginomycetes

            Class Hymenomycetes

 

LICHENS

    Biology

    Lichen compounds

    Evolution

    Classification

        Historical background

        Diagnostic characters

        Orders of Ascomycetes with lichen-forming fungi

        Orders of Basidiomycetes with lichen-forming fungi

        "Imperfect lichenized fungi"

    Geographical distribution

    Ecology

    Declining lichens

        Pollution

        Loss of habitat

    Future investigations

 

SYSTEMATIC BOTANY

    Aims and Philosophy of Plant Systematics

        The Role of Plant Classifications in Life Support Systems

        Philosophical Basis of Plant Classification

    History and Development of Plant Systematics

The First Empirical Plant Classificatory Systems

The Herbals

The First Plant Taxonomists

Natural Plant Classifications

Phylogenetic Classifications

Modern Phylogenetic Plant Classifications

Plant Molecular Systematics

    Plant Diversity

Is There a Plant Kingdom?

Evolution of Photosynthesis

Endosymbiotic Origin of Plastids

The Plant Kingdom

The First Land Plants

Plants with Vascular Tissue

The First Terrestrial Ecosystems

The Lycopod-line

The Sphenofern-line

The First Forests

The Evolution of Seeds

The First Seed Plants

The Pine-line

Ancient Mesozoic Floras

The Anthophyte Clade

Evolution of the Flower and Flowering Plants

Why Did Angiosperms Evolve?

What Did the First Angiosperms Look Like?

When Did the Angiosperms Evolve?

Where Did the Angiosperms Evolve?

From What Gymnosperm Ancestor Did the Angiosperms Evolve?

Flowering Plants

 

ALGAE

Cyanobacteria

Glaucophyta

Rhodophyta

Chlorophyta

Heterokontophyta

Dinophyta

Haptophyta

Cryptophyta

Euglenophyta

 

BRYOPHYTE SYSTEMATICS

Morphology

Anthocerotae (Hornworts)

Hepaticae (Liverworts)

Musci (Mosses)

Distinguishing Features Among Groups

Sources of Systematic Data

Ontogeny

Cytology

Chemistry

Phylogenetic Relationships

Classification

Future Research

 

SYSTEMATIC ZOOLOGY: INVERTEBRATES

    Animal phylogeny: data sources and interpretation

Morphology

Molecules

Fossils

    Systematic zoology of the Metazoa

Porifera, Placozoa, Cnidaria, Ctenophora

Platyhelminthes, Nemertea

Mollusca, Sipunculida, Entoprocta, Cycliophora

Annelida, Pogonophora, Myzostomida, Echiura

Onychophora, Tardigrada, Arthropoda

Rotifera, Acanthocephala, Gnathostomulida, Micrognathozoa

Priapulida, Kinorhyncha, Loricifera

Nematoda, Nematomorpha, Gastrotricha, Chaetognatha

Ectoprocta, Phoronida, Brachiopoda

Urochordata, Cephalochordata

Enteropneusta, Pterobranchia, Echinodermata

Some extant "problematica"

    Alternative hypotheses of metazoan relationships

Morphology

Molecules

    Reconciliation and pathways to future progress

 

PORIFERA, CNIDARIA, AND CTENOPHORA

    Porifera

General Remarks

Importance

Form and Function

Size and Shape

Color, Consistency, and Surface

Water Current System

Cell Types

Skeleton

Life Cycle

Sexual Reproduction

Embryonic Development and Larvae

Asexual Reproduction

Regeneration, Individuality, and Life Span

Ecology

Evolution and Paleontology

Phylogeny and Systematics

    Cnidaria

General Remarks

Importance

Form and Function

Body Structure

Size

Color

Cell Types

Life Cycle

Sexual Reproduction and Development

Asexual Reproduction

Coloniality and Polymorphism

Metagenetic Cycles

Ecology

Phylogeny and Systematics

    Ctenophora

 

PLATYHELMINTHES, NEMERTEA, AND “ASCHELMINTHES”

General Morphology

Platyhelminthes, the Flatworms

Nemertea (Nemertini), the Ribbon Worms

 “Aschelminthes”

Gnathifera

Gnathostomulida

Micrognathozoa (Limnognathia Maerski)

Rotifera

Acanthocephala

Cycliophora (Symbion Pandora)

Nemathelminthes

Gastrotricha

Nematoda, the Roundworms

Nematomorpha, the Horsehair Worms

Priapulida

Kinorhyncha

Loricifera

 

ANNELIDA

    Basic annelid organization

Coelom

Metamerism

Chaetae

    Annelid morphology

Body wall

Parapodia

Nervous system

Sense organs

Circulation and respiratory structures

Segmental organs

    Annelid diversity

Polychaeta

Clitellata

Unusual or controversial groups placed in Annelida

Echiura

Myzostomida

Siboglinidae (= Pogonophora and Vestimentifera)

    Annelid behaviour and physiology

Locomotion

Feeding

Osmoregulation

Symbiosis

    Annelid reproduction and development

Asexual reproduction in Polychaeta

Sexual reproduction in Polychaeta

Sexual Reproduction in Clitellata

    Annelid phylogeny

Fossil Annelida

 

ARTHROPODS OTHER THAN INSECTS

    Myriapoda

Chilopoda

Diplopoda

Symphyla

Pauropoda

General aspects of myriapod biology

    Crustacea

Remipedia

Cephalocarida

Branchiopoda

Maxillopoda

Malacostraca

Phyllocarida

Hoplocarida

Eumalacostraca

General aspects of crustacean biology

    Cheliceriformes [= Cheliceromorpha]

Pycnogonida

Chelicerata

Merostomata

Scorpiones

Pseudoscorpiones

Solifugae

Opiliones

Palpigradi

Amblypygi

Uropygi

Schizomida

Araneae

Ricinulei

Acari

General aspects of cheliceriform biology

 

INSECTS AND OTHER HEXAPODOUS ARTHROPODS

What are Insects?

Why are Insects so Successful?

Insect Life Cycles

Ecological Importance of Insects

Insect Diversity

Superclass: Hexapoda - Hexapods, Insects sensu lato (i.e., in the broad sense)

Class: Entognatha - Entognathans (may be a paraphyletic group)

Class: Insecta sensu stricto (= Ectognatha) - True Insects

 

 

MOLLUSCA

Relationships and Higher Classification

General Morphology of the Mollusca

Gastropoda

Major Groups, Diversity and Adaptive Radiation

    Bivalvia (=Pelecypoda)

Major Groups, Diversity and Adaptive Radiation

    Cephalopoda

Major Groups, Diversity and Adaptive Radiation

    Other Classes Living and Extinct

Polyplacophora

"Aplacophora" (Caudofoveata and Solenogastres)

Monoplacophora (Helcionelloidea and Tergomya)

Scaphopoda

Extinct Groups that have been Suggested as Additional Classes

Current Knowledge

Geographic Diversity

Ecological Diversity

Conservation

Marine

Freshwater

Terrestrial

    Economic Importance

Bivalvia

Cephalopoda

Gastropoda

 

ECHINODERMATA

    General Morphology of the Echinodermata

Recognizing an Echinoderm

Organization of the Echinoderm Body

Morphological Pattern

Embryological Pattern

Classification and Relationships of the Echinodermata

Edrioasteroid-like Echinoderms

Morphology of Edrioasteroid-like Echinoderms

Paleoecology of Edrioasteroid-like Echinoderms

    Blastozoans

Morphology of Blastozoans

Paleoecology of Blastozoans

    Crinoidea

Morphology of Crinoids

Reproduction and Development of Crinoids

Ecology and Behavior of Crinoids

Economic Importance of Crinoids

    Asteroidea

Morphology of Asteroids

Reproduction and Development of Asteroids

Ecology and Behavior of Asteroids

Economic Importance of Asteroids

    Ophiuroidea

Morphology of Ophiuroids

Reproduction and Development of Ophiuroids

Ecology and Behavior of Ophiuroids

Economic Importance of Ophiuroids

Holothuroidea

Morphology of Holothuroids

Reproduction and Development of Holothuroids

Ecology and Behavior of Holothuroids

Economic Importance of Holothuroids

Echinoidea

Morphology of Echinoids

Reproduction and Development of Echinoids

Ecology and Behavior of Echinoids

Economic Importance of Echinoids

Ongoing and Future Research

 

OTHER INVERTEBRATE TAXA

Mesozoa

Rhombozoa

Orthonectida

Xenoturbellida (Xenoturbella Bocki)

Kamptozoa (Entoprocta)

Sipunculida

Tardigrada, the Water-bears

Onychophora, the Velvet Worms

Chaetognatha, the Arrow Worms

Tentaculata, or Lophophorata

Phoronida

Bryozoa (Ectoprocta)

Brachiopoda

Hemichordata

Pterobranchia

Enteropneusta

 

TUNICATA AND CEPHALOCHORDATA

Tunicata

Ascidiacea

Tunic

Morphology and Physiology

Reproduction and Development

Larval Morphology and Metamorphosis

Ecology and Economic Relevance

Systematics

Thaliacea

Pyrosomes

Salps

Doliolids

Appendicularia

Morphology and Behavior

Relevance in Marine Ecosystems

    Cephalochordata

Morphology and Physiology

Reproduction and Development

 

PISCES

The Biology of Fishes

    Respiration

Osmoregulation

Buoyancy

Prey Detection and Predator Avoidance

Reproduction

The Emergence of Fishes

Agnatha (Jawless Fishes)

Gnathostomata (Jawed Fishes)

Placodermi (Armored Fishes)

Acanthodii (Spiny-Finned Fishes)

Chondrichthyes (Cartilaginous Fishes)

Sarcopterygii (Lobe-Finned Fishes)

Actinopterygii (Ray-Finned Fishes)

Basal (Pre-Teleostean) Actinopterygians

The Teleostei

 

AMPHIBIA

Classification of Amphibia

Gymnophiona

Morphology

Life Cycle

Distribution

Feeding and Predators

Little-known Caecilians

Caudata

Morphology

Life Cycle

Distribution

Adult Variation

Feeding

Larval Variation

Anura

Morphology

Life Cycle

Variations in Life-history Strategies

Distribution

Adult Variation

Tadpole Variation

Food

Vocalization

Sound Production

Hearing

Metamorphosis

Ecological Role

Declining Amphibians

Implicated Factors

Ultraviolet Radiation

Loss of Habitat

Introduced Predators

Pollutants

Fungus

Long-term Drought

Human Consumption

Skin Secretions

Future Investigations

 

REPTILES

Diversity and Systematics

A Brief Overview of the Families of Extant Reptiles

The Role of Reptiles in Ecosystems

Humans and Reptiles

The Global Conservation Status of Reptiles

Threatening Processes

 

BIRDS

Basic attributes

Body

Feathers

Power production

Song

Life Span

Structure and Physiology

Outer shape

Integument

Feathers

Scales and Claws

Bill Covering

Wings

Legs

Tail

Feeding Apparatus

Bill

Mouth and Tongue

Digestive System

Urinary System.

Respiratory System.

Circulatory System.

Reproductive System.

Sensory System

Hearing

Sight

Smell.

Brain.  

Temperature Regulation

Yearly Cycle

Establishing Territories

Courtship

Mating Systems

Nests

Eggs

Incubation

Hatching

Care of the young

Molt

Fall Migration

Direction Finding

Navigation

Winter

Spring

Flocking Behavior

Behavior

Distribution

Evolution and Classification

Origin and fossils

Classification.

Class Aves

Subclass Sauriurae (all extinct)

Subclass Ornithurae

Superorder Neornithae

 

MAMMALS

Mammal characteristics

Hair

Epidermal glands

Lactation and parental care

Cardiovascular and respiratory specializations

Feeding specializations

Improved auditory acuity

An enlarged brain

Mammal evolution 

Ancestors

Evolutionary trends during therapsid-mammal transition

The origin of mammals

Early mammals

Systematics   

Traditional classification

Molecules remodel the eutherian phylogenetic tree

Modern mammal classification

Diversity and conservation priorities  

Diversity

Global threats and conservation priorities

Future research

 

PRIMATE EVOLUTION

Archaic Primates

Early Euprimates“Primates of Modern Aspect”

Adapiforms and Omomyiforms

Early Simians

Later Simian Radiations

Early Monkeys

The Epoch of the Apes

The Origins of Modern Primate G

Nonhuman Primates

The Human Lineage

Early Hominins

The Genus “Homo”